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CARMA3 Is Critical for the Initiation of Allergic Airway Inflammation
Journal article   Peer reviewed

CARMA3 Is Critical for the Initiation of Allergic Airway Inflammation

Benjamin Causton, Ravisankar A Ramadas, Josalyn L Cho, Khristianna Jones, Ana Pardo-Saganta, Jayaraj Rajagopal, Ramnik J Xavier and Benjamin D Medoff
The Journal of immunology (1950), Vol.195(2), pp.683-694
07/15/2015
DOI: 10.4049/jimmunol.1402983
PMCID: PMC4489191
PMID: 26041536

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Abstract

Innate immune responses to allergens by airway epithelial cells (AECs) help initiate and propagate the adaptive immune response associated with allergic airway inflammation in asthma. Activation of the transcription factor NF-κB in AECs by allergens or secondary mediators via G protein-coupled receptors (GPCRs) is an important component of this multifaceted inflammatory cascade. Members of the caspase recruitment domain family of proteins display tissue-specific expression and help mediate NF-κB activity in response to numerous stimuli. We have previously shown that caspase recruitment domain-containing membrane-associated guanylate kinase protein (CARMA)3 is specifically expressed in AECs and mediates NF-κB activation in these cells in response to stimulation with the GPCR agonist lysophosphatidic acid. In this study, we demonstrate that reduced levels of CARMA3 in normal human bronchial epithelial cells decreases the production of proasthmatic mediators in response to a panel of asthma-relevant GPCR ligands such as lysophosphatidic acid, adenosine triphosphate, and allergens that activate GPCRs such as Alternaria alternata and house dust mite. We then show that genetically modified mice with CARMA3-deficient AECs have reduced airway eosinophilia and proinflammatory cytokine production in a murine model of allergic airway inflammation. Additionally, we demonstrate that these mice have impaired dendritic cell maturation in the lung and that dendritic cells from mice with CARMA3-deficient AECs have impaired Ag processing. In conclusion, we show that AEC CARMA3 helps mediate allergic airway inflammation, and that CARMA3 is a critical signaling molecule bridging the innate and adaptive immune responses in the lung.
Ovalbumin - immunology Dendritic Cells - immunology Epithelial Cells - drug effects NF-kappa B - immunology CARD Signaling Adaptor Proteins - deficiency Dendritic Cells - pathology Male CARD Signaling Adaptor Proteins - genetics Adenosine Triphosphate - pharmacology Allergens - immunology Asthma - immunology Lysophospholipids - pharmacology Dendritic Cells - drug effects Female Cytokines - immunology Adaptive Immunity CARD Signaling Adaptor Proteins - immunology Lung - pathology Signal Transduction Mice, Inbred C57BL Cells, Cultured Gene Expression Regulation Epithelial Cells - pathology Mice, Transgenic Immunity, Innate Asthma - chemically induced Asthma - genetics Alternaria - immunology Animals NF-kappa B - genetics Epithelial Cells - immunology Lung - drug effects Mice Asthma - pathology Cytokines - biosynthesis Lung - immunology Pyroglyphidae - immunology

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