Logo image
Constitutive Nuclear Localization of NFAT in Foxp3(+) Regulatory T Cells Independent of Calcineurin Activity
Journal article   Peer reviewed

Constitutive Nuclear Localization of NFAT in Foxp3(+) Regulatory T Cells Independent of Calcineurin Activity

Qiuxia Li, Arvind Shakya, Xiaohua Guo, Hongbo Zhang, Dean Tantin, Peter E. Jensen and Xinjian Chen
The Journal of immunology (1950), Vol.188(9), pp.4268-4277
05/01/2012
DOI: 10.4049/jimmunol.1102376
PMID: 22490438

View Online

Abstract

Foxp3 plays an essential role in conferring suppressive functionality to CD4(+)/Foxp3(+) regulatory T cells (Tregs). Although studies showed that Foxp3 has to form cooperative complexes with NFAT to bind to target genes, it remains unclear whether NFAT is available in the nucleus of primary Tregs for Foxp3 access. It is generally believed that NFAT in resting cells resides in the cytoplasm, and its nuclear translocation depends on calcineurin (CN) activation. We report that a fraction of NFAT protein constitutively localizes in the nucleus of primary Tregs, where it selectively binds to Foxp3 target genes. Treating Tregs with CN inhibitor does not induce export of NFAT from the nucleus, indicating that its nuclear translocation is independent of CN activity. Consistently, Tregs are resistant to CN inhibitors in the presence of IL-2 and continue to proliferate in response to anti-CD3 stimulation, whereas proliferation of non-Tregs is abrogated by CN inhibitors. In addition, PMA, which activates other transcription factors required for T cell activation but not NFAT, selectively induces Treg proliferation in the absence of ionomycin. TCR interaction with self-MHC class II is not required for PMA-induced Treg proliferation. Tregs expanded by PMA or in the presence of CN inhibitors maintain Treg phenotype and functionality. These findings shed light on Treg biology, paving the way for strategies to selectively activate Tregs. The Journal of Immunology, 2012, 188: 4268-4277.
Immunology Life Sciences & Biomedicine Science & Technology

Details

Metrics

Logo image