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Functional analysis of fruitless gene expression by transgenic manipulations of Drosophila courtship
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Functional analysis of fruitless gene expression by transgenic manipulations of Drosophila courtship

Adriana Villella, Sarah L. Ferri, Jonathan D. Krystal and Jeffrey C. Hall
Proceedings of the National Academy of Sciences - PNAS, Vol.102(46), pp.16550-16557
Inaugural Article
09/22/2005
DOI: 10.1073/pnas.0507056102
PMCID: PMC1225315
PMID: 16179386
url
https://doi.org/10.1073/pnas.0507056102View
Published (Version of record) Open Access

Abstract

A gal4 -containing enhancer–trap called C309 was previously shown to cause subnormal courtship of Drosophila males toward females and courtship among males when driving a conditional disrupter of synaptic transmission ( shi TS ). We extended these manipulations to analyze all features of male-specific behavior, including courtship song, which was almost eliminated by driving shi TS at high temperature. In the context of singing defects and homosexual courtship affected by mutations in the fru gene, a tra -regulated component of the sex-determination hierarchy, we found a C309 / tra F combination also to induce high levels of courtship between pairs of males and “chaining” behavior in groups; however, these doubly transgenic males sang normally. Because production of male-specific FRU M protein is regulated by TRA, we hypothesized that a fru -derived transgene encoding the male ( M ) form of an I nhibitory R NA ( fru MIR ) would mimic the effects of tra F ; but C309 / fru MIR males exhibited no courtship chaining, although they courted other males in single-pair tests. Double-labeling of neurons in which GFP was driven by C309 revealed that 10 of the 20 CNS clusters containing FRU M in wild-type males included coexpressing neurons. Histological analysis of the developing CNS could not rationalize the absence of tra F or fru MIR effects on courtship song, because we found C309 to be coexpressed with FRU M within the same 10 neuronal clusters in pupae. Thus, we hypothesize that elimination of singing behavior by the C309 / shi TS combination involves neurons acting downstream of FRU M cells
Biological Sciences C309 enhancer trap inhibitory fru RNA transgene reproductive behavior shiTS transgene traF transgene

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