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Head Rotation or Dissociation?: A Study of Exponential Rate Processes in Chemically Skinned Rabbit Muscle Fibers When MgATP Concentration Is Changed
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Head Rotation or Dissociation?: A Study of Exponential Rate Processes in Chemically Skinned Rabbit Muscle Fibers When MgATP Concentration Is Changed

Masataka Kawai
Biophysical journal, Vol.22(1), pp.97-103
04/1978
DOI: 10.1016/S0006-3495(78)85473-3
PMCID: PMC1473409
PMID: 638228
url
https://doi.org/10.1016/S0006-3495(78)85473-3View
Published (Version of record) Open Access

Abstract

The mechanical response of fully activated muscle bundles (one to five fibers) to sinusoidal length perturbation (∼0.4% L 0 ) was studied as a function of MgATP concentration. The frequency response (0.25-167 Hz; corresponding to 1 ms time resolution) of chemically skinned rabbit muscle fibers was resolved into three exponential rate processes, (A), (B), and (C). At 20°C, the apparent rate constants associated with the fast exponential lead (2π c = 388-588 s -1 ) and the oscillatory work (2π b = 59-116 s -1 ) both increase with increment of the MgATP concentration from 1 to 5 mM, and they both saturate for further increase. Over the whole range of MgATP concentrations the slow exponential lead (2π a = 9-7 s -1 ) remains constant. The effect of MgATP on processes (B) and (C) can be interpreted in the context of the biochemical evidence, in which MgATP enters the cross-bridge cycle after the desorption of the product, and the binding of MgATP to rigorlike cross-bridges promotes a rapid dissociation of actomyosin (Lymn and Taylor, 1971. Biochemistry. 10: 4617-4624.). The effect is not predicted by a model for force generation in which head rotation dominates the fast component (“stage 2” of Huxley and Simmons, 1971. Nature (Lond.). 233: 533-538. and 1973. Cold Spring Harbor Symp. Quant. Biol. 37: 669-680.), and head dissociation dominates the slow component (“phase 4” of Huxley, 1974. J. Physiol. (Lond.). 243: 1-43; Julian et al., 1974. Biophys. J. 14: 546-562.).

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