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Intestinal Neurod1 expression impairs paneth cell differentiation and promotes enteroendocrine lineage specification
Journal article   Open access   Peer reviewed

Intestinal Neurod1 expression impairs paneth cell differentiation and promotes enteroendocrine lineage specification

Hui Joyce Li, Subir K Ray, Ning Pan, Jody Haigh, Bernd Fritzsch and Andrew B Leiter
Scientific reports, Vol.9(1), pp.19489-11
12/20/2019
DOI: 10.1038/s41598-019-55292-7
PMCID: PMC6925293
PMID: 31862906
url
https://doi.org/10.1038/s41598-019-55292-7View
Published (Version of record) Open Access

Abstract

Transcription factor Neurod1 is required for enteroendocrine progenitor differentiation and maturation. Several earlier studies indicated that ectopic expression of Neurod1 converted non- neuronal cells into neurons. However, the functional consequence of ectopic Neurod1 expression has not been examined in the GI tract, and it is not known whether Neurod1 can similarly switch cell fates in the intestine. We generated a mouse line that would enable us to conditionally express Neurod1 in intestinal epithelial cells at different stages of differentiation. Forced expression of Neurod1 throughout intestinal epithelium increased the number of EECs as well as the expression of EE specific transcription factors and hormones. Furthermore, we observed a substantial reduction of Paneth cell marker expression, although the expressions of enterocyte-, tuft- and goblet-cell specific markers are largely not affected. Our earlier study indicated that Neurog3+ progenitor cells give rise to not only EECs but also Goblet and Paneth cells. Here we show that the conditional expression of Neurod1 restricts Neurog3+ progenitors to adopt Paneth cell fate, and promotes more pronounced EE cell differentiation, while such effects are not seen in more differentiated Neurod1+ cells. Together, our data suggest that forced expression of Neurod1 programs intestinal epithelial cells more towards an EE cell fate at the expense of the Paneth cell lineage and the effect ceases as cells mature to EE cells.
Paneth Cells - metabolism Intestinal Mucosa - metabolism Basic Helix-Loop-Helix Transcription Factors - genetics Enterocytes - metabolism Gene Expression Regulation - genetics Mice, Inbred C57BL Gene Expression Regulation - physiology Goblet Cells - metabolism Blotting, Western Nerve Tissue Proteins - metabolism Cell Differentiation - genetics Animals Basic Helix-Loop-Helix Transcription Factors - metabolism Fluorescent Antibody Technique Mice Cell Differentiation - physiology

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