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Local Impermeant Anions Establish the Neuronal Chloride Concentration
Journal article   Peer reviewed

Local Impermeant Anions Establish the Neuronal Chloride Concentration

J Glykys, V Dzhala, K Egawa, T Balena, Y Saponjian, K V Kuchibhotla, B J Bacskai, K T Kahle, T Zeuthen and K J Staley
Science (American Association for the Advancement of Science), Vol.343(6171), pp.670-675
02/07/2014
DOI: 10.1126/science.1245423
PMCID: PMC4220679
PMID: 24503855
url
http://doi.org/10.1126/science.1245423View
Open Access

Abstract

Neuronal intracellular chloride concentration [Cl(-)](i) is an important determinant of γ-aminobutyric acid type A (GABA(A)) receptor (GABA(A)R)-mediated inhibition and cytoplasmic volume regulation. Equilibrative cation-chloride cotransporters (CCCs) move Cl(-) across the membrane, but accumulating evidence suggests factors other than the bulk concentrations of transported ions determine [Cl(-)](i). Measurement of [Cl(-)](i) in murine brain slice preparations expressing the transgenic fluorophore Clomeleon demonstrated that cytoplasmic impermeant anions ([A](i)) and polyanionic extracellular matrix glycoproteins ([A](o)) constrain the local [Cl(-)]. CCC inhibition had modest effects on [Cl(-)](i) and neuronal volume, but substantial changes were produced by alterations of the balance between [A](i) and [A](o). Therefore, CCCs are important elements of Cl(-) homeostasis, but local impermeant anions determine the homeostatic set point for [Cl(-)], and hence, neuronal volume and the polarity of local GABA(A)R signaling.

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