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Mitochondrial-targeted catalase is good for the old mouse proteome, but not for the young: 'reverse' antagonistic pleiotropy?
Journal article   Open access   Peer reviewed

Mitochondrial-targeted catalase is good for the old mouse proteome, but not for the young: 'reverse' antagonistic pleiotropy?

Nathan Basisty, Dao-Fu Dai, Arni Gagnidze, Lemuel Gitari, Jeanne Fredrickson, Yvonne Maina, Richard P Beyer, Mary J Emond, Edward J Hsieh, Michael J MacCoss, …
Aging cell, Vol.15(4), pp.634-645
08/2016
DOI: 10.1111/acel.12472
PMCID: PMC4933659
PMID: 27061426
url
https://doi.org/10.1111/acel.12472View
Published (Version of record) Open Access

Abstract

Reactive oxygen species (ROS) are highly reactive oxygen-containing molecules associated with aging and a broad spectrum of pathologies. We have previously shown that transgenic expression of the antioxidant enzyme catalase targeted to the mitochondria (mCAT) in mice reduces ROS, attenuates age-related disease, and increases lifespan. However, it has been increasingly recognized that ROS also has beneficial roles in signaling, hormesis, stress response, and immunity. We therefore hypothesized that mCAT might be beneficial only when ROS approaches pathological levels in older age and might not be advantageous at a younger age when basal ROS is low. We analyzed abundance and turnover of the global proteome in hearts and livers of young (4 month) and old (20 month) mCAT and wild-type (WT) mice. In old hearts and livers of WT mice, protein half-lives were reduced compared to young, while in mCAT mice the reverse was observed; the longest half-lives were seen in old mCAT mice and the shortest in young mCAT. Protein abundance of old mCAT hearts recapitulated a more youthful proteomic expression profile (P-value < 0.01). However, young mCAT mice partially phenocopied the older wild-type proteome (P-value < 0.01). Age strongly interacts with mCAT, consistent with antagonistic pleiotropy in the reverse of the typical direction. These findings underscore the contrasting roles of ROS in young vs. old mice and indicate the need for better understanding of the interaction between dose and age in assessing the efficacy of therapeutic interventions in aging, including mitochondrial antioxidants.
Biomarkers - metabolism Catalase - metabolism Animals Metabolic Networks and Pathways Liver - metabolism Mice, Inbred C57BL Myocardium - metabolism Half-Life Genetic Pleiotropy Mitochondria - metabolism Proteome - metabolism Aging - metabolism

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