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Novel function of the Fanconi anemia group J or RECQ1 helicase to disrupt protein-DNA complexes in a replication protein A-stimulated manner
Journal article   Open access   Peer reviewed

Novel function of the Fanconi anemia group J or RECQ1 helicase to disrupt protein-DNA complexes in a replication protein A-stimulated manner

Joshua A Sommers, Taraswi Banerjee, Twila Hinds, Bingbing Wan, Marc S Wold, Ming Lei and Robert M Brosh Jr
The Journal of biological chemistry, Vol.289(29), pp.19928-19941
07/18/2014
DOI: 10.1074/jbc.M113.542456
PMCID: PMC4106313
PMID: 24895130
url
https://doi.org/10.1074/jbc.M113.542456View
Published (Version of record) Open Access

Abstract

Understanding how cellular machinery deals with chromosomal genome complexity is an important question because protein bound to DNA may affect various cellular processes of nucleic acid metabolism. DNA helicases are at the forefront of such processes, yet there is only limited knowledge how they remodel protein-DNA complexes and how these mechanisms are regulated. We have determined that representative human RecQ and Fe-S cluster DNA helicases are potently blocked by a protein-DNA interaction. The Fanconi anemia group J (FANCJ) helicase partners with the single-stranded DNA-binding protein replication protein A (RPA) to displace BamHI-E111A bound to duplex DNA in a specific manner. Protein displacement was dependent on the ATPase-driven function of the helicase and unique properties of RPA. Further biochemical studies demonstrated that the shelterin proteins TRF1 and TRF2, which preferentially bind the telomeric repeat found at chromosome ends, effectively block FANCJ from unwinding the forked duplex telomeric substrate. RPA, but not the Escherichia coli single-stranded DNA-binding protein or shelterin factor Pot1, stimulated FANCJ ejection of TRF1 from the telomeric DNA substrate. FANCJ was also able to displace TRF2 from the telomeric substrate in an RPA-dependent manner. The stimulation of helicase-catalyzed protein displacement is also observed with the DNA helicase RECQ1, suggesting a conserved functional interaction of RPA-interacting helicases. These findings suggest that partnerships between RPA and interacting human DNA helicases may greatly enhance their ability to dislodge proteins bound to duplex DNA, an activity that is likely to be highly relevant to their biological roles in DNA metabolism.
Recombinant Proteins - metabolism Basic-Leucine Zipper Transcription Factors - metabolism RecQ Helicases - metabolism Humans Werner Syndrome Helicase Deoxyribonuclease BamHI - metabolism Substrate Specificity Telomeric Repeat Binding Protein 1 - metabolism Recombinant Proteins - genetics Replication Protein A - metabolism DNA - metabolism DNA - genetics DNA - chemistry Fanconi Anemia Complementation Group Proteins - metabolism Macromolecular Substances - chemistry Base Sequence Replication Protein A - genetics Exodeoxyribonucleases - metabolism Protein Binding Nucleic Acid Conformation Macromolecular Substances - metabolism Amino Acid Substitution

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