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The Eukaryotic Two-Component Histidine Kinase Sln1p Regulates OCH1 via the Transcription Factor, Skn7p
Journal article   Open access   Peer reviewed

The Eukaryotic Two-Component Histidine Kinase Sln1p Regulates OCH1 via the Transcription Factor, Skn7p

Sheng Li, Susan Dean, Zhijian Li, Joe Horecka, Robert J Deschenes and Jan S Fassler
Molecular biology of the cell, Vol.13(2), pp.412-424
02/2002
DOI: 10.1091/mbc.01-09-0434
PMCID: PMC65637
PMID: 11854400
url
https://doi.org/10.1091/mbc.01-09-0434View
Published (Version of record) Open Access

Abstract

The yeast “two-component” osmotic stress phosphorelay consists of the histidine kinase, Sln1p, the phosphorelay intermediate, Ypd1p and two response regulators, Ssk1p and Skn7p, whose activities are regulated by phosphorylation of a conserved aspartyl residue in the receiver domain. Dephospho-Ssk1p leads to activation of the hyper-osmotic response (HOG) pathway, whereas phospho-Skn7p presumably leads to activation of hypo-osmotic response genes. The multifunctional Skn7 protein is important in oxidative as well as osmotic stress; however, the Skn7p receiver domain aspartate that is the phosphoacceptor in the SLN1 pathway is dispensable for oxidative stress. Like many well-characterized bacterial response regulators, Skn7p is a transcription factor. In this report we investigate the role of Skn7p in osmotic response gene activation. Our studies reveal that the Skn7p HSF-like DNA binding domain interacts with a cis -acting element identified upstream of OCH1 that is distinct from the previously defined HSE-like Skn7p binding site. Our data support a model in which Skn7p receiver domain phosphorylation affects transcriptional activation rather than DNA binding to this class of DNA binding site.

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