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Metabolic control of smooth muscle cell phenotype switching in atherosclerosis
Preprint   Open access

Metabolic control of smooth muscle cell phenotype switching in atherosclerosis

Rong-Mo Zhang, Xiaolong Zhu, Hosung Bae, Jiasheng Zhang, Yanming Li, Pei-Yu Chen, Ying H. Shen, George Tellides, Nathaniel Snyder, Cholsoon Jang, …
bioRxiv
Cold Spring Harbor Laboratory, 1.1
05/21/2026
DOI: 10.64898/2026.05.19.726223
PMID: 42239337
url
https://doi.org/10.64898/2026.05.19.726223View
Preprint (Author's original) This preprint has not been evaluated by subject experts through peer review. Preprints may undergo extensive changes and/or become peer-reviewed journal articles. Open Access

Abstract

The loss of smooth muscle cell (SMC) contractile phenotype contributes to various diseases including atherosclerosis. However, its metabolic basis is not entirely elucidated. Since the transforming growth factor beta (TGFβ) signaling is among principal regulators of SMC contractility, we studied metabolic regulation of TGFβ signaling in SMCs in vitro and atherosclerotic mouse models and human lesions. We found that TGFβ induced Ac-CoA synthetase 2 (ACSS2)-dependent Ac-CoA production, by suppressing pyruvate dehydrogenase kinase 4 (PDK4). This stabilized R-SMADs and TGFβ receptor 1, preserving SMC contractile phenotype. SMC-specific PDK4 knockout mimicked the effect of TGFβ signaling both metabolically and phenotypically, increasing glucose-derived synthesis of Ac-CoA and SMC contractile phenotype. SMC-specific Pdk4 knockout in ApoE knockout mice reduced atherosclerosis. Furthermore, human specimens demonstrated a strong correlation between PDK4 level and atherosclerosis severity. These findings indicate that continuous TGFβ signaling, critical to the maintenance of the normal SMC contractile state and is regulated by PDK4 and carbohydrate metabolism. Reducing PDK4 metabolically restricts aortic plaque growth via TGFβ-dependent SMC contractility.
Cell Biology

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